Uncle King Julien

Uncle King Julien XII (voiced by Henry Winkler and Dee Bradley Baker) is the main antagonist of his nephew's own story, the hidden main antagonist of All Hail King Julien and the secondary antagonist of All Hail King Julien: Exile. He is rude, dishonest, obnoxious, evil, lazy, sarcastic, deceiving, selfish, vain, cowardly and idiotic.

Towards the end of the series, Uncle gives up on taking the throne back, because he fell in love with Zora and wanted them to run away together. Julien pardons them both and allows them to marry and live a happy life together. This explains why he didn't try to take the throne back from Julien when he, Maurice, and Mort left Madagascar for good in the second movie.